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J Pharmacol Exp Ther 263:964970 Kannan R, Mittur A, Bao Y, Tsuruo T, Kaplowitz N (1999) GSH transport in immortalized mouse brain endothelial cells: evidence for apical localization of a sodium-dependent GSH transporter

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Nuclear glutathione

In that paper and in their 2013 paper, they showed the following: D-lactate dehydrogenase is able to donate electrons to a synthetic electron acceptor added experimentally only when mitochondria are turned inside out, not when they are kept intact, which indicates that the enzyme is associated with the inner face of the inner mitochondrial membrane, facing the matrix

5.3 Addressing translational challenges Despite these promising results, several challenges must be addressed to optimize SFNs clinical application